Species distribution models (SDMs), which aim to predict species occurrence based on environmental variables, are widely used to monitor and respond to biodiversity change. Recent deep learning advances for SDMs have been shown to perform well on complex and heterogeneous datasets, but their effectiveness remains limited by spatial biases in the data. In this paper, we revisit deep SDMs from a Bayesian perspective and introduce BATIS, a novel and practical framework wherein prior predictions are updated iteratively using limited observational data. Models must appropriately capture both aleatoric and epistemic uncertainty to effectively combine fine-grained local insights with broader ecological patterns. We benchmark an extensive set of uncertainty quantification approaches on a novel dataset including citizen science observations from the eBird platform. Our empirical study shows how Bayesian deep learning approaches can greatly improve the reliability of SDMs in data-scarce locations, which can contribute to ecological understanding and conservation efforts.

Traditional methods for species distribution models (SDMs) use environmental data to predict the distribution of species across geographic space.

Species distribution models (SDMs) are widely used to predict species' geographic distributions, serving as critical tools for ecological research and conservation planning. Typically, SDMs relate species occurrences to environmental variables representing abiotic factors, such as temperature, precipitation, and soil properties. However, species distributions are also strongly influenced by biotic interactions with other species, which are often overlooked. While some methods partially address this limitation by incorporating biotic interactions, they often assume symmetrical pairwise relationships between species and require consistent co-occurrence data. In practice, species observations are sparse, and the availability of information about the presence or absence of other species varies significantly across locations. To address these challenges, we propose CISO, a deep learning-based method for species distribution modeling Conditioned on Incomplete Species Observations. CISO enables predictions to be conditioned on a flexible number of species observations alongside environmental variables, accommodating the variability and incompleteness of available biotic data. We demonstrate our approach using three datasets representing different species groups: sPlotOpen for plants, SatBird for birds, and a new dataset, SatButterfly, for butterflies. Our results show that including partial biotic information improves predictive performance on spatially separate test sets. When conditioned on a subset of species within the same dataset, CISO outperforms alternative methods in predicting the distribution of the remaining species. Furthermore, we show that combining observations from multiple datasets can improve performance. CISO is a promising ecological tool, capable of incorporating incomplete biotic information and identifying potential interactions between species from disparate taxa.

Climate change poses an extreme threat to biodiversity, making it imperative to efficiently model the geographical range of different species. The availability of large-scale remote sensing images and environmental data has facilitated the use of machine learning in Species Distribution Models (SDMs), which aim to predict the presence of a species at any given location. Traditional SDMs, reliant on expert observation, are labor-intensive, but advancements in remote sensing and citizen science data have facilitated machine learning approaches to SDM development. However, these models often struggle with leveraging spatial relationships between different inputs -- for instance, learning how climate data should inform the data present in satellite imagery -- without upsampling or distorting the original inputs. Additionally, location information and ecological characteristics at a location play a crucial role in predicting species distribution models, but these aspects have not yet been incorporated into state-of-the-art approaches. In this work, we introduce MiTREE: a multi-input Vision-Transformer-based model with an ecoregion encoder. MiTREE computes spatial cross-modal relationships without upsampling as well as integrates location and ecological context. We evaluate our model on the SatBird Summer and Winter datasets, the goal of which is to predict bird species encounter rates, and we find that our approach improves upon state-of-the-art baselines.

To address the interlinked biodiversity and climate crises, we need an understanding of where species occur and how these patterns are changing. However, observational data on most species remains very limited, and the amount of data available varies greatly between taxonomic groups. We introduce the problem of predicting species occurrence patterns given (a) satellite imagery, and (b) known information on the occurrence of other species. To evaluate algorithms on this task, we introduce SatButterfly, a dataset of satellite images, environmental data and observational data for butterflies, which is designed to pair with the existing SatBird dataset of bird observational data. To address this task, we propose a general model, R-Tran, for predicting species occurrence patterns that enables the use of partial observational data wherever found. We find that R-Tran outperforms other methods in predicting species encounter rates with partial information both within a taxon (birds) and across taxa (birds and butterflies). Our approach opens new perspectives to leveraging insights from species with abundant data to other species with scarce data, by modelling the ecosystems in which they co-occur.